KRIGOLSON TEACHING
  • NEUROSCIENCE
    • NEUROSCIENCE 100 >
      • NEURO 100 INTRODUCTION
      • NEURO 101 ADVANCED
      • NEURO 102 AGING
      • NEURO 103 MEMORY
      • NEURO 104 DECISION MAKING
      • NEURO 105 LEARNING
      • Research Statistics
    • NRSC 500B / MEDS 470
  • Kinesiology
    • EPHE 245 >
      • LABORATORY
      • PRACTICUM
    • EPHE 357
  • STATISTICS
    • LECTURE >
      • INTRODUCTION TO R
      • DESCRIPTIVE STATISTICS
      • VISUALIZING DATA
      • Correlation and Regression
      • MULTIPLE REGRESSION
      • LOGIC OF NHST
      • T TESTS
      • ANOVA
      • POST HOC ANALYSIS
      • NON PARAMETRIC STATISTICS
      • FACTORIAL ANOVA
      • Repeated Measures ANOVA
      • Mixed ANOVA
      • MULTIVARIATE ANOVA
      • THE NEW STATISTICS
      • Bayesian Methods
    • ASSIGNMENTS >
      • Introduction to R >
        • INTRODUCTION TO R
        • LOADING DATA
        • DATA TABLES
      • Descriptive Statistics >
        • Mean, Median, and Mode
        • VARIANCE
        • CONFIDENCE INTERVALS
        • SHORTCUTS
      • Visualizing Data >
        • PLOTTING BASICS
        • BAR GRAPHS
        • BOXPLOTS
        • HISTOGRAMS
        • USING GGPLOT I
        • USING GGPLOT II
        • USING GGPLOT III
      • Correlation and Regression >
        • CORRELATION
        • REGRESSION
      • MULTIPLE REGRESSION >
        • MULTIPLE REGRESSION
      • Logic of NHST >
        • Sample Size and Variance
        • DISTRIBUTIONS
        • TESTING DISTRIBUTIONS
      • T-Tests >
        • Single Sample TTests
        • Paired Sample TTests
        • Independent Sample TTests
      • ANOVA >
        • ANOVA ASSUMPTIONS
        • ANOVA
      • POST HOC ANALYSIS >
        • POSTHOC ANALYSIS
      • NON PARAMETRIC STATISTICS >
        • WILCOXON TEST
        • WILCOXON SIGNED TEST
        • MULTIPLE GROUPS
      • FACTORIAL ANOVA
      • REPEATED MEASURES ANOVA >
        • RM ANOVA
        • TREND ANALYSIS
      • MIXED ANOVA
      • MULTIVARIATE ANOVA
      • THE NEW STATISTICS
      • BAYESIAN TTESTS
    • RESOURCES
    • R TIPS
  • Directed Studies
    • Advanced Topics in Motor Control A
    • Advanced Topics in Motor Control B
    • An Introduction to EEG
    • Advanced EEG and ERP Methods
    • Neural Correlates of Human Reward Processing
    • Independent Research Project
  • MATLAB
    • THE BASICS >
      • Hello World
      • BASIC MATHEMATICS
      • VARIABLES
      • Matrices
      • Writing Scripts
      • PATHS AND DIRECTORIES
      • USER INPUT
      • FOR LOOPS
      • WHILE LOOPS
      • IF STATEMENTS
      • RANDOM NUMBERS
    • STATISTICS >
      • LOADING DATA
      • DESCRIPTIVE STATISTICS
      • MAKING FUNCTIONS
      • BAR GRAPHS
      • LINE GRAPHS
      • TTESTS
    • EXPERIMENTS: THE BASICS >
      • DRAWING A CIRCLE
      • DRAWING MULTIPLE OBJECTS
      • DRAWING TEXT
      • DRAWING AN IMAGE
      • PLAYING A TONE
      • KEYBOARD INPUT
      • BUILDING A TRIAL
      • BUILDING TRIALS
      • NESTED LOOPS
      • RIGHT OR WRONG
      • SAVING DATA
    • EXPERIMENTS: ADVANCED >
      • STROOP
      • N BACK
      • Oddball
      • Animation
      • VIDEO
    • EEG and ERP Analysis >
      • ERP Analysis
  • RESOURCES
    • EXCEL
    • HOW TO READ A RESEARCH PAPER
    • HOW TO WRITE A RESEARCH PAPER
  • Workshops
    • Iowa State EEG Workshop 2018
  • Python
    • The Basics >
      • Setting Up Python
      • Hello, world!
      • Basic Math & Using Import
      • Variables
      • Matrices
      • Scripts
      • User Input
      • For Loops

8. LEARNING

 We learn in two principle ways, through repetition and through feedback.

The theoretical basis for learning through repetition is Hebb's postulate - "neurons that fire together, wire together." In terms of what happens, repetition is thought to first induce long term potentiation (LTP). Following the creation of an LTP like state, it is believed that long term changes occur at the synaptic level.

Preparatory Questions

1. What is LTP? Use the following two papers as references: Caporale 2008 and Lomo 2003.

2. What are the long term changes that occur at the synapse related to learning through repetition. Use the following two papers as a reference: Luscher 2012 and Mayford 2012. 

3. Put what you have learned together. When an action is repeated, how does learning occur? This CHAPTER may help you.

We also learn through feedback. Intriguingly, it appears that feedback dependent learning follows the principles of reinforcement learning. Specifically, when we experience outcomes we compute prediction errors - the difference between our expectancies and our outcomes and we use these to modify behaviour.

Preparatory Questions

Use the first chapter of Sutton 2018 to answer the following three questions.

4. What is reinforcement learning?

5. What is a prediction error?

6. How are prediction errors used to modify behaviour?

7. In terms of how prediction errors are reflected within the brain, it is thought that dopamine plays a prominent role. Review Schultz et al. 1997. What is the key finding and how does it related to how reinforcement learning might occur in the human brain? (focus on the role of dopamine).

8. Studies using fMRI have provided insight into the regions of the brain involved in feedback dependent learning. What are the roles of the striatum, medial prefrontal cortex, and anterior cingulate cortex in the feedback learning process? THIS review paper should help.

9. How does it all fit together? In other words, what is one model for a neural feedback dependent learning system involving the prefrontal cortex, the ACC, the striatum, and the midbrain dopamine system. THIS paper should help, but note that this is only one of many theories.

EXAM QUESTION

8. How do we learn? Make sure you discuss learning via repetition and learning via feedback. Ensure you discuss both the theoretical and neural aspects of learning.





  • NEUROSCIENCE
    • NEUROSCIENCE 100 >
      • NEURO 100 INTRODUCTION
      • NEURO 101 ADVANCED
      • NEURO 102 AGING
      • NEURO 103 MEMORY
      • NEURO 104 DECISION MAKING
      • NEURO 105 LEARNING
      • Research Statistics
    • NRSC 500B / MEDS 470
  • Kinesiology
    • EPHE 245 >
      • LABORATORY
      • PRACTICUM
    • EPHE 357
  • STATISTICS
    • LECTURE >
      • INTRODUCTION TO R
      • DESCRIPTIVE STATISTICS
      • VISUALIZING DATA
      • Correlation and Regression
      • MULTIPLE REGRESSION
      • LOGIC OF NHST
      • T TESTS
      • ANOVA
      • POST HOC ANALYSIS
      • NON PARAMETRIC STATISTICS
      • FACTORIAL ANOVA
      • Repeated Measures ANOVA
      • Mixed ANOVA
      • MULTIVARIATE ANOVA
      • THE NEW STATISTICS
      • Bayesian Methods
    • ASSIGNMENTS >
      • Introduction to R >
        • INTRODUCTION TO R
        • LOADING DATA
        • DATA TABLES
      • Descriptive Statistics >
        • Mean, Median, and Mode
        • VARIANCE
        • CONFIDENCE INTERVALS
        • SHORTCUTS
      • Visualizing Data >
        • PLOTTING BASICS
        • BAR GRAPHS
        • BOXPLOTS
        • HISTOGRAMS
        • USING GGPLOT I
        • USING GGPLOT II
        • USING GGPLOT III
      • Correlation and Regression >
        • CORRELATION
        • REGRESSION
      • MULTIPLE REGRESSION >
        • MULTIPLE REGRESSION
      • Logic of NHST >
        • Sample Size and Variance
        • DISTRIBUTIONS
        • TESTING DISTRIBUTIONS
      • T-Tests >
        • Single Sample TTests
        • Paired Sample TTests
        • Independent Sample TTests
      • ANOVA >
        • ANOVA ASSUMPTIONS
        • ANOVA
      • POST HOC ANALYSIS >
        • POSTHOC ANALYSIS
      • NON PARAMETRIC STATISTICS >
        • WILCOXON TEST
        • WILCOXON SIGNED TEST
        • MULTIPLE GROUPS
      • FACTORIAL ANOVA
      • REPEATED MEASURES ANOVA >
        • RM ANOVA
        • TREND ANALYSIS
      • MIXED ANOVA
      • MULTIVARIATE ANOVA
      • THE NEW STATISTICS
      • BAYESIAN TTESTS
    • RESOURCES
    • R TIPS
  • Directed Studies
    • Advanced Topics in Motor Control A
    • Advanced Topics in Motor Control B
    • An Introduction to EEG
    • Advanced EEG and ERP Methods
    • Neural Correlates of Human Reward Processing
    • Independent Research Project
  • MATLAB
    • THE BASICS >
      • Hello World
      • BASIC MATHEMATICS
      • VARIABLES
      • Matrices
      • Writing Scripts
      • PATHS AND DIRECTORIES
      • USER INPUT
      • FOR LOOPS
      • WHILE LOOPS
      • IF STATEMENTS
      • RANDOM NUMBERS
    • STATISTICS >
      • LOADING DATA
      • DESCRIPTIVE STATISTICS
      • MAKING FUNCTIONS
      • BAR GRAPHS
      • LINE GRAPHS
      • TTESTS
    • EXPERIMENTS: THE BASICS >
      • DRAWING A CIRCLE
      • DRAWING MULTIPLE OBJECTS
      • DRAWING TEXT
      • DRAWING AN IMAGE
      • PLAYING A TONE
      • KEYBOARD INPUT
      • BUILDING A TRIAL
      • BUILDING TRIALS
      • NESTED LOOPS
      • RIGHT OR WRONG
      • SAVING DATA
    • EXPERIMENTS: ADVANCED >
      • STROOP
      • N BACK
      • Oddball
      • Animation
      • VIDEO
    • EEG and ERP Analysis >
      • ERP Analysis
  • RESOURCES
    • EXCEL
    • HOW TO READ A RESEARCH PAPER
    • HOW TO WRITE A RESEARCH PAPER
  • Workshops
    • Iowa State EEG Workshop 2018
  • Python
    • The Basics >
      • Setting Up Python
      • Hello, world!
      • Basic Math & Using Import
      • Variables
      • Matrices
      • Scripts
      • User Input
      • For Loops